About the Authors

Ceiridwen J. Edwards

Contributed equally to this work with: Ceiridwen J. Edwards, David A. Magee, Stephen D. E. Park

Current address: Research Laboratory for Archaeology, Oxford University, Oxford, United Kingdom

Affiliation Smurfit Institute of Genetics, Trinity College, Dublin, Ireland

David A. Magee

Contributed equally to this work with: Ceiridwen J. Edwards, David A. Magee, Stephen D. E. Park

Affiliation Animal Genomics Laboratory, School of Agriculture, Food Science and Veterinary Medicine, College of Life Sciences, University College Dublin, Dublin, Ireland

Stephen D. E. Park

Contributed equally to this work with: Ceiridwen J. Edwards, David A. Magee, Stephen D. E. Park

Affiliation Animal Genomics Laboratory, School of Agriculture, Food Science and Veterinary Medicine, College of Life Sciences, University College Dublin, Dublin, Ireland

Paul A. McGettigan

Affiliation Animal Genomics Laboratory, School of Agriculture, Food Science and Veterinary Medicine, College of Life Sciences, University College Dublin, Dublin, Ireland

Amanda J. Lohan

Affiliation Conway Institute of Biomolecular and Biomedical Research, University College Dublin, Dublin, Ireland

Alison Murphy

Affiliation Conway Institute of Biomolecular and Biomedical Research, University College Dublin, Dublin, Ireland

Emma K. Finlay

Affiliation Smurfit Institute of Genetics, Trinity College, Dublin, Ireland

Beth Shapiro

Current address: Department of Biology, Pennsylvania State University, University Park, Pennsylvania, United States of America

Affiliation Henry Wellcome Ancient Biomolecules Centre, Department of Zoology, Oxford University, Oxford, United Kingdom

Andrew T. Chamberlain

Affiliation Department of Archaeology, University of Sheffield, Sheffield, United Kingdom

Martin B. Richards

Affiliation Institute of Integrative and Comparative Biology, Faculty of Biological Sciences, University of Leeds, Leeds, United Kingdom

Daniel G. Bradley

Affiliation Smurfit Institute of Genetics, Trinity College, Dublin, Ireland

Brendan J. Loftus

Affiliation Conway Institute of Biomolecular and Biomedical Research, University College Dublin, Dublin, Ireland

David E. MacHugh

david.machugh@ucd.ie

Affiliations Animal Genomics Laboratory, School of Agriculture, Food Science and Veterinary Medicine, College of Life Sciences, University College Dublin, Dublin, Ireland, Conway Institute of Biomolecular and Biomedical Research, University College Dublin, Dublin, Ireland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CJE DAM DGB BL DM. Performed the experiments: CJE DAM AJL AM BS. Analyzed the data: CJE DAM SDEP PAM EKF MR DGB DM. Contributed reagents/materials/analysis tools: AJL AM BS AC BL. Wrote the paper: CJE DAM SDEP DGB DM.