About the Authors

He-Ge Chen

Contributed equally to this work with: He-Ge Chen, Wen-Jun Han, Mi Deng, Jichao Qin

Affiliations Department of Biochemistry & Molecular Biology, College of Medicine, University of Nebraska Medical Center, Omaha, Nebraska, United States of America, Key Laboratory of Protein Chemistry and Developmental Biology of Education Ministry of China, College of Life Sciences, Hunan Normal University, Changsha, Hunan, China

Wen-Jun Han

Contributed equally to this work with: He-Ge Chen, Wen-Jun Han, Mi Deng, Jichao Qin

Affiliation Key Laboratory of Protein Chemistry and Developmental Biology of Education Ministry of China, College of Life Sciences, Hunan Normal University, Changsha, Hunan, China

Mi Deng

Contributed equally to this work with: He-Ge Chen, Wen-Jun Han, Mi Deng, Jichao Qin

Affiliation Department of Biochemistry & Molecular Biology, College of Medicine, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Jichao Qin

Contributed equally to this work with: He-Ge Chen, Wen-Jun Han, Mi Deng, Jichao Qin

Affiliation Department of Biochemistry & Molecular Biology, College of Medicine, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Dan Yuan

Affiliations Department of Biochemistry & Molecular Biology, College of Medicine, University of Nebraska Medical Center, Omaha, Nebraska, United States of America, Key Laboratory of Protein Chemistry and Developmental Biology of Education Ministry of China, College of Life Sciences, Hunan Normal University, Changsha, Hunan, China

Jin-Ping Liu

Affiliation Department of Biochemistry & Molecular Biology, College of Medicine, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Ling Xiao

Affiliations Department of Biochemistry & Molecular Biology, College of Medicine, University of Nebraska Medical Center, Omaha, Nebraska, United States of America, Key Laboratory of Protein Chemistry and Developmental Biology of Education Ministry of China, College of Life Sciences, Hunan Normal University, Changsha, Hunan, China

Lili Gong

Affiliation Department of Biochemistry & Molecular Biology, College of Medicine, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Songping Liang

Affiliation Key Laboratory of Protein Chemistry and Developmental Biology of Education Ministry of China, College of Life Sciences, Hunan Normal University, Changsha, Hunan, China

Jian Zhang

Affiliation Key Laboratory of Protein Chemistry and Developmental Biology of Education Ministry of China, College of Life Sciences, Hunan Normal University, Changsha, Hunan, China

Yun Liu

Affiliation Key Laboratory of Protein Chemistry and Developmental Biology of Education Ministry of China, College of Life Sciences, Hunan Normal University, Changsha, Hunan, China

David Wan-Cheng Li

dwli1688@hotmail.com

Affiliations Department of Biochemistry & Molecular Biology, College of Medicine, University of Nebraska Medical Center, Omaha, Nebraska, United States of America, Key Laboratory of Protein Chemistry and Developmental Biology of Education Ministry of China, College of Life Sciences, Hunan Normal University, Changsha, Hunan, China, Department of Ophthalmology & Visual Sciences, College of Medicine, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HGC WJH MD JQ JZ YL DWL. Performed the experiments: HGC WJH MD JQ DY JPL LX LG. Analyzed the data: HGC WJH MD JQ DY JPL LX LG SL DWL. Contributed reagents/materials/analysis tools: SL JZ YL. Wrote the paper: DWL.