About the Authors

Caroline L. Benn

Affiliations Department of Medical and Molecular Genetics, King's College London School of Medicine, King's College London, London, United Kingdom, Pfizer Regenerative Medicine, UCB Granta Park, Great Abington, Cambridge, United Kingdom

Rachel Butler

Affiliations Department of Medical and Molecular Genetics, King's College London School of Medicine, King's College London, London, United Kingdom, Nuffield Laboratory of Ophthalmology, University of Oxford, The John Radcliffe Hospital, Oxford, United Kingdom

Lydia Mariner

Affiliation Department of Medical and Molecular Genetics, King's College London School of Medicine, King's College London, London, United Kingdom

Jude Nixon

Affiliation Department of Medical and Molecular Genetics, King's College London School of Medicine, King's College London, London, United Kingdom

Hilary Moffitt

Affiliation Department of Medical and Molecular Genetics, King's College London School of Medicine, King's College London, London, United Kingdom

Michal Mielcarek

Affiliation Department of Medical and Molecular Genetics, King's College London School of Medicine, King's College London, London, United Kingdom

Ben Woodman

Affiliation Department of Medical and Molecular Genetics, King's College London School of Medicine, King's College London, London, United Kingdom

Gillian P. Bates

gillian.bates@kcl.ac.uk

Affiliation Department of Medical and Molecular Genetics, King's College London School of Medicine, King's College London, London, United Kingdom

Competing Interests

The R6/2 mice are licensed by King's College London for commercial use.

Author Contributions

Conceived and designed the experiments: GB. Performed the experiments: CLB RB LM HM MM. Analyzed the data: CLB RB LM JN MM. Contributed reagents/materials/analysis tools: BW. Wrote the paper: CLB GB.