About the Authors

Bronwen Martin

Affiliation Laboratory of Neurosciences, National Institute on Aging Intramural Research Program, Biomedical Research Center, Baltimore, Maryland, United States of America

Randall Brenneman

Affiliation Laboratory of Neurosciences, National Institute on Aging Intramural Research Program, Biomedical Research Center, Baltimore, Maryland, United States of America

Kevin G. Becker

Affiliation Research Resources Branch, National Institute on Aging Intramural Research Program, Biomedical Research Center, Baltimore, Maryland, United States of America

Marjan Gucek

Affiliation Mass Spectrometry/Proteomics Facility at Johns Hopkins School of Medicine, Baltimore, Maryland, United States of America

Robert N. Cole

Affiliation Mass Spectrometry/Proteomics Facility at Johns Hopkins School of Medicine, Baltimore, Maryland, United States of America

Stuart Maudsley

maudsleyst@grc.nia.nih.gov

Affiliation Laboratory of Neurosciences, National Institute on Aging Intramural Research Program, Biomedical Research Center, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SM. Performed the experiments: BM RB MG RNC SM. Analyzed the data: BM SM. Contributed reagents/materials/analysis tools: KGB RNC SM. Wrote the paper: BM SM.