About the Authors

Hyun-Gwan Lee

Affiliation Department of Biology, Huck Institute Genetics Graduate Program, Pennsylvania State University, University Park, Pennsylvania, United States of America

Young-Cho Kim

Affiliation Department of Biology, Huck Institute Neuroscience Graduate Program, Pennsylvania State University, University Park, Pennsylvania, United States of America

Jennifer S. Dunning

Affiliation Department of Biology, Pennsylvania State University, University Park, Pennsylvania, United States of America

Kyung-An Han

To whom correspondence should be addressed. E-mail: kxh29@psu.edu

Affiliations Department of Biology, Huck Institute Genetics Graduate Program, Pennsylvania State University, University Park, Pennsylvania, United States of America, Department of Biology, Huck Institute Neuroscience Graduate Program, Pennsylvania State University, University Park, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KH HL. Performed the experiments: KH HL YK JD. Analyzed the data: KH HL YK. Contributed reagents/materials/analysis tools: HL YK. Wrote the paper: KH.