About the Authors

Ching-Lin Hsieh

Affiliation Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Eric Chang

Affiliation Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Andrew Tseng

Affiliation Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Christopher Ptak

Affiliation Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Li-Chen Wu

Affiliation Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Chun-Li Su

Affiliation Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Sean P. McDonough

Affiliation Department of Biomedical Science, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Yi-Pin Lin

Affiliation Division of Infectious Disease, Wadsworth Center, New York State Department of Health, Albany, New York, United States of America

Yung-Fu Chang

yc42@cornell.edu

Affiliation Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CLH CP YFC. Performed the experiments: CLH EC AT LCW CLS SPM. Analyzed the data: CLH CP YPL YFC. Wrote the paper: CLH CP SPM YFC.