About the Authors

Carine Brouat

carine.brouat@ird.fr

Affiliation IRD, UMR022 CBGP, Montferrier-sur-Lez, France

Soanandrasana Rahelinirina

Affiliation Institut Pasteur de Madagascar, Antananarivo, Madagascar

Anne Loiseau

Affiliation INRA, UMR1062 CBGP, Montferrier-sur-Lez, France

Lila Rahalison

Address Current address: Centers for Disease Control and Prevention, Atlanta, Georgia, United States of America.

Affiliation Institut Pasteur de Madagascar, Antananarivo, Madagascar

Minoariso Rajerison

Affiliation Institut Pasteur de Madagascar, Antananarivo, Madagascar

Dominique Laffly

Affiliations Département de Géographie, UFR Sciences Espaces Sociétés, Université de Toulouse, Toulouse, France, GEODE UMR 5602 CNRS, Toulouse, France

Pascal Handschumacher

Affiliation IRD, UMR912 SE4S, Marseille, France

Jean-Marc Duplantier

Affiliation IRD, UMR022 CBGP, Montferrier-sur-Lez, France

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JMD LR CB. Performed the experiments: JMD SR AL. Analyzed the data: CB SR DL. Contributed reagents/materials/analysis tools: MR PH. Wrote the paper: CB.