About the Authors

Jonathan D. Stoltzfus

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Samuel Minot

Affiliation Department of Microbiology, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Matthew Berriman

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Thomas J. Nolan

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

James B. Lok

jlok@vet.upenn.edu

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JDS SM JBL. Performed the experiments: JDS. Analyzed the data: JDS. Contributed reagents/materials/analysis tools: JDS SM MB TJN. Wrote the paper: JDS.