About the Authors

Kazumasa Yokoyama

Affiliation Division of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Hokkaido, Japan

Hyun Kim

Affiliation Division of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Hokkaido, Japan

Tetsu Mukai

Affiliation Leprosy Research Center, National Institute of Infectious Diseases, Higashimurayama, Tokyo, Japan

Masanori Matsuoka

Affiliation Leprosy Research Center, National Institute of Infectious Diseases, Higashimurayama, Tokyo, Japan

Chie Nakajima

Affiliation Division of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Hokkaido, Japan

Yasuhiko Suzuki

suzuki@czc.hokudai.ac.jp

Affiliations Division of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Hokkaido, Japan, JST/JICA-SATREPS, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TM MM CN YS. Performed the experiments: KY HK CN. Analyzed the data: KY CN YS. Contributed reagents/materials/analysis tools: KY TM MM. Wrote the paper: KY TM MM CN YS.