About the Authors

Yi-Chin Fan

Affiliation Graduate Institute of Microbiology and Public Health, College of Veterinary Medicine, National Chung Hsing University, Taichung, Taiwan

Jo-Mei Chen

Affiliation Graduate Institute of Microbiology and Public Health, College of Veterinary Medicine, National Chung Hsing University, Taichung, Taiwan

Hsien-Chung Chiu

Affiliation Department of Periodontology, School of Dentistry, National Defense Medical Center and Tri-Service General Hospital, Taipei, Taiwan

Yi-Ying Chen

Affiliation Graduate Institute of Microbiology and Public Health, College of Veterinary Medicine, National Chung Hsing University, Taichung, Taiwan

Jen-Wei Lin

Affiliation Graduate Institute of Microbiology and Public Health, College of Veterinary Medicine, National Chung Hsing University, Taichung, Taiwan

Chen-Chang Shih

Affiliation Department of Neurology, Mennonite Christian Hospital, Hualien, Taiwan

Chih-Ming Chen

Affiliations Graduate Institute of Microbiology and Public Health, College of Veterinary Medicine, National Chung Hsing University, Taichung, Taiwan, Division of Infectious Disease, Department of Internal Medicine, Tungs' Taichung MetroHarbor Hospital, Taichung, Taiwan

Chao-Chin Chang

Affiliation Graduate Institute of Microbiology and Public Health, College of Veterinary Medicine, National Chung Hsing University, Taichung, Taiwan

Gwong-Jen J. Chang

Affiliation Arboviral Diseases Branch, Center for Disease Control and Prevention, Fort Collins, Colorado, United States of America

Shyan-Song Chiou

sschiou@dragon.nchu.edu.tw

Affiliation Graduate Institute of Microbiology and Public Health, College of Veterinary Medicine, National Chung Hsing University, Taichung, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YCF SSC. Performed the experiments: YCF JMC YYC JWL. Analyzed the data: YCF CCC GJJC SSC. Contributed reagents/materials/analysis tools: HCC CCS CMC. Wrote the paper: YCF SSC.