About the Authors

Zhisheng Dang

Affiliations Division of Collaboration and Education, Research Center for Zoonosis Control, Hokkaido University, Sapporo, Hokkaido, Japan, State Key Laboratory of Veterinary Etiological Biology, Key Laboratory of Grazing Animal Diseases MOA, Key Laboratory of Veterinary Parasitology of Gansu Province, Lanzhou Veterinary Research Institute, Chinese Academy of Agricultural Sciences, Gansu, People's Republic of China

Kinpei Yagi

Affiliation Department of Biological Science, Hokkaido Institute of Public Health, Sapporo, Hokkaido, Japan

Yuzaburo Oku

Affiliation Parasitology Laboratory, School of Veterinary Medicine, Faculty of Agriculture, Tottori University, Tottori, Japan

Hirokazu Kouguchi

Affiliation Department of Biological Science, Hokkaido Institute of Public Health, Sapporo, Hokkaido, Japan

Kiichi Kajino

Affiliation Division of Collaboration and Education, Research Center for Zoonosis Control, Hokkaido University, Sapporo, Hokkaido, Japan

Jun Matsumoto

Affiliation Laboratory of Medical Zoology, Nihon University College of Bioresource Sciences, Fujisawa, Japan

Ryo Nakao

Affiliation Division of Collaboration and Education, Research Center for Zoonosis Control, Hokkaido University, Sapporo, Hokkaido, Japan

Hiroyuki Wakaguri

Affiliation Department of Medical Genome Science, Graduate School of Frontier Science, The University of Tokyo, Tokyo, Japan

Atsushi Toyoda

Affiliations RIKEN Genomic Sciences Center, Yokohama, Kanagawa, Japan, Comparative Genomics Laboratory, National Institute of Genetics, Mishima, Shizuoka, Japan

Hong Yin

Affiliation State Key Laboratory of Veterinary Etiological Biology, Key Laboratory of Grazing Animal Diseases MOA, Key Laboratory of Veterinary Parasitology of Gansu Province, Lanzhou Veterinary Research Institute, Chinese Academy of Agricultural Sciences, Gansu, People's Republic of China

Chihiro Sugimoto

sugimoto@czc.hokudai.ac.jp

Affiliation Division of Collaboration and Education, Research Center for Zoonosis Control, Hokkaido University, Sapporo, Hokkaido, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZD KY YO KK CS. Performed the experiments: ZD KY HK RN. Analyzed the data: ZD YO HY CS. Contributed reagents/materials/analysis tools: KY JM HW AT. Wrote the paper: ZD YO JM CS.