About the Authors

Steve J. Torr, Glyn A. Vale
Natural Resources Institute, University of Greenwich, London, United Kingdom

Glyn A. Vale
South African Centre for Epidemiological Modelling and Analysis (SACEMA), University of Stellenbosch, Stellenbosch, South Africa

Corresponding Author

s.torr@greenwich.ac.uk

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GAV SJT. Performed the experiments: GAV. Analyzed the data: SJT GAV. Wrote the paper: SJT GAV.