About the Authors

David T. S. Hayman, Nicholas Johnson, Daniel L. Horton, Philip R. Wakeley, Ashley C. Banyard, Anthony R. Fooks
Rabies and Wildlife Zoonoses Group, Veterinary Laboratories Agency – Weybridge, Surrey, United Kingdom

David T. S. Hayman
Cambridge Infectious Diseases Consortium, Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

David T. S. Hayman
Institute of Zoology, Zoological Society of London, Regent's Park, London, United Kingdom

Jessica Hedge
Institute for Evolutionary Biology, University of Edinburgh, Edinburgh, United Kingdom

Shoufeng Zhang
Veterinary Services Laboratory, Ministry of Food and Agriculture, Accra, Ghana

Andy Alhassan
Changchun Veterinary Research Institute, Chinese Academy of Agricultural Science, Changchun, Jilin, People's Republic of China

Anthony R. Fooks
The National Centre for Zoonosis Research, University of Liverpool, Leahurst, Neston, United Kingdom

Corresponding Author

t.fooks@vla.defra.gsi.gov.uk

Competing Interests

Prof. Tony Fooks is an Academic Editor for PLoS ONE.

Author Contributions

Conceived and designed the experiments: DTSH PRW AA ARF. Performed the experiments: DTSH NJ DLH PRW ACB SZ. Analyzed the data: DTSH NJ DLH JH ACB SZ. Contributed reagents/materials/analysis tools: PRW AA ARF. Wrote the paper: DTSH NJ ARF.