About the Authors

Li-Qun Fang

Contributed equally to this work with: Li-Qun Fang, Xian-Jun Wang, Song Liang

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Xian-Jun Wang

Contributed equally to this work with: Li-Qun Fang, Xian-Jun Wang, Song Liang

Affiliation Shandong Center for Disease Control and Prevention, Jinan, China

Song Liang

Contributed equally to this work with: Li-Qun Fang, Xian-Jun Wang, Song Liang

Affiliation College of Public Health, The Ohio State University, Columbus, Ohio, United States of America

Yan-Li Li

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Shao-Xia Song

Affiliation Shandong Center for Disease Control and Prevention, Jinan, China

Wen-Yi Zhang

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Quan Qian

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Ya-Pin Li

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Lan Wei

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Zhi-Qiang Wang

Affiliation Shandong Center for Disease Control and Prevention, Jinan, China

Hong Yang

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Wu-Chun Cao

caowc@nic.bmi.ac.cn

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LQF XJW SL WCC. Analyzed the data: LQF XJW SL YLL SXS WYZ QQ YPL LW ZQW HY WCC. Contributed reagents/materials/analysis tools: LQF XJW YLL SXS WYZ QQ YPL LW ZQW HY. Wrote the paper: LQF XJW SL WCC.