About the Authors

Kathryn J. Ray

Affiliation F.I. Proctor Foundation, University of California, San Francisco, California, United States of America

Thomas M. Lietman

tom.lietman@ucsf.edu

Affiliations F.I. Proctor Foundation, University of California, San Francisco, California, United States of America, Department of Ophthalmology, University of California, San Francisco, California, United States of America, Department of Epidemiology & Biostatistics, University of California, San Francisco, California, United States of America, Institute for Global Health, University of California, San Francisco, California, United States of America

Travis C. Porco

Affiliations F.I. Proctor Foundation, University of California, San Francisco, California, United States of America, Department of Ophthalmology, University of California, San Francisco, California, United States of America, Department of Epidemiology & Biostatistics, University of California, San Francisco, California, United States of America

Jeremy D. Keenan

Affiliations F.I. Proctor Foundation, University of California, San Francisco, California, United States of America, Department of Ophthalmology, University of California, San Francisco, California, United States of America

Robin L. Bailey

Affiliation Department of Infectious and Tropical Diseases, London School of Hygiene and Tropical Medicine, London, United Kingdom

Anthony W. Solomon

Affiliation Department of Infectious and Tropical Diseases, London School of Hygiene and Tropical Medicine, London, United Kingdom

Matthew J. Burton

Affiliation Department of Infectious and Tropical Diseases, London School of Hygiene and Tropical Medicine, London, United Kingdom

Emma Harding-Esch

Affiliation Department of Infectious and Tropical Diseases, London School of Hygiene and Tropical Medicine, London, United Kingdom

Martin J. Holland

Affiliation Department of Infectious and Tropical Diseases, London School of Hygiene and Tropical Medicine, London, United Kingdom

David Mabey

Affiliation Department of Infectious and Tropical Diseases, London School of Hygiene and Tropical Medicine, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TML TCP RLB AWS MJB DM. Performed the experiments: KJR TML. Analyzed the data: KJR TML JDK. Contributed reagents/materials/analysis tools: KJR TML TCP JDK RLB MJB EHE MJH DM. Wrote the paper: KJR TML TCP JDK AWS DM.