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Contributed equally to this work with:
Gareth J Hughes
¤ Current address: Veterinary Epidemiology Research Unit, SAC (Scottish Agricultural College), Inverness, Scotland
Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, Scotland
Chelsea and Westminster Hospital, London, United Kingdom
*To whom correspondence should be addressed. E-mail: A.Leigh-Brown@ed.ac.uk
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