About the Authors

Michael N Weedon

Affiliation Department of Diabetes Research and Vascular Medicine, Peninsula Medical School, Exeter, United Kingdom

Mark I McCarthy

Affiliation Oxford Centre for Diabetes, Endocrinology, and Metabolism, Churchill Hospital, Headington, Oxford, United Kingdom

Graham Hitman

Affiliation Centre of Diabetes and Metabolic Medicine, Bart's and the London Queen Mary's School of Medicine and Dentistry, University of London, London, United Kingdom

Mark Walker

Affiliation Department of Medicine, School of Medicine, Newcastle-upon-Tyne, United Kingdom

Christopher J Groves

Affiliation Oxford Centre for Diabetes, Endocrinology, and Metabolism, Churchill Hospital, Headington, Oxford, United Kingdom

Eleftheria Zeggini

Affiliation Oxford Centre for Diabetes, Endocrinology, and Metabolism, Churchill Hospital, Headington, Oxford, United Kingdom

N. William Rayner

Affiliation Oxford Centre for Diabetes, Endocrinology, and Metabolism, Churchill Hospital, Headington, Oxford, United Kingdom

Beverley Shields

Affiliation Department of Diabetes Research and Vascular Medicine, Peninsula Medical School, Exeter, United Kingdom

Katharine R Owen

Affiliation Department of Diabetes Research and Vascular Medicine, Peninsula Medical School, Exeter, United Kingdom

Andrew T Hattersley

Affiliation Department of Diabetes Research and Vascular Medicine, Peninsula Medical School, Exeter, United Kingdom

Timothy M Frayling

To whom correspondence should be addressed. E-mail: Tim.Frayling@pms.ac.uk

Affiliation Department of Diabetes Research and Vascular Medicine, Peninsula Medical School, Exeter, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MNW, MIM, CJG, EZ, NWR, ATH, and TMF were involved in the design and implementation of the study. MNW, MIM, CJG, EZ, NWR, ATH, and TMF were involved in the writing up of the study. MIM, GH, MW, BS, KRO, and ATH were involved in the design and execution of studies to collect DNA samples. All authors contributed to and read the final manuscript.