About the Authors

Ben S Cooper

To whom correspondence should be addressed. E-mail: ben.cooper@hpa.org.uk

Affiliation Statistics, Modelling, and Bioinformatics Department, Centre for Infections, Health Protection Agency, London, United Kingdom

Richard J Pitman

Affiliation Statistics, Modelling, and Bioinformatics Department, Centre for Infections, Health Protection Agency, London, United Kingdom

W. John Edmunds

Affiliation Statistics, Modelling, and Bioinformatics Department, Centre for Infections, Health Protection Agency, London, United Kingdom

Nigel J Gay

Affiliation Statistics, Modelling, and Bioinformatics Department, Centre for Infections, Health Protection Agency, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

BSC, WJE, and NJG conceived the study. RJP obtained and processed the travel data. BSC implemented and analysed the model. All authors contributed to writing the manuscript.