Center for Human Genetics, Duke University Medical Center, Durham, North Carolina, United States of America,
Department of Biostatistics and Bioinformatics, Duke University Medical Center, Durham, North Carolina, United States of America
Centre for Eye Research Australia, University of Melbourne, Royal Victorian Eye and Ear Hospital, Melbourne, Australia,
Lions Eye Institute, University of Western Australia, Centre for Ophthalmology and Visual Science, Perth, Australia
¶These authors also contributed equally to this work.