About the Authors

Masatoshi Mutazono

Affiliation Department of Biological Sciences, Graduate School of Science and Technology, Kumamoto University, Kurokami, Chuo-ku, Kumamoto, Japan

Misato Morita

Affiliation Department of Biological Sciences, Graduate School of Science and Technology, Kumamoto University, Kurokami, Chuo-ku, Kumamoto, Japan

Chihiro Tsukahara

Affiliation Department of Biological Sciences, Graduate School of Science and Technology, Kumamoto University, Kurokami, Chuo-ku, Kumamoto, Japan

Madoka Chinen

Current address: API Co., Ltd., Kanohsakurada-cho, Gifu, Japan.

Affiliation Department of Biological Sciences, Graduate School of Science and Technology, Kumamoto University, Kurokami, Chuo-ku, Kumamoto, Japan

Shiori Nishioka

Affiliation Department of Biological Sciences, Graduate School of Science and Technology, Kumamoto University, Kurokami, Chuo-ku, Kumamoto, Japan

Tatsuhiro Yumikake

Affiliation Department of Biological Sciences, Graduate School of Science and Technology, Kumamoto University, Kurokami, Chuo-ku, Kumamoto, Japan

Kohei Dohke

Affiliation Laboratory of Chromosome Function and Regulation, Graduate School of Frontier Biosciences, Osaka University, Osaka, Japan

Misuzu Sakamoto

Affiliation Department of Biological Sciences, Graduate School of Science and Technology, Kumamoto University, Kurokami, Chuo-ku, Kumamoto, Japan

Takashi Ideue

Affiliation Department of Biological Sciences, Graduate School of Science and Technology, Kumamoto University, Kurokami, Chuo-ku, Kumamoto, Japan

Jun-ichi Nakayama

Affiliations Graduate School of Natural Sciences, Nagoya City University, Nagoya, Japan, Division of Chromatin Regulation, National Institute for Basic Biology, Okazaki, Japan

Kojiro Ishii

Affiliation Laboratory of Chromosome Function and Regulation, Graduate School of Frontier Biosciences, Osaka University, Osaka, Japan

Tokio Tani

ttani@sci.kumamoto-u.ac.jp

Affiliation Department of Biological Sciences, Graduate School of Science and Technology, Kumamoto University, Kurokami, Chuo-ku, Kumamoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: TT MMu. Data curation: MMu TT. Formal analysis: MMu. Funding acquisition: TT MMu. Investigation: MMu MMo CT MC SN TY MS KD KI TI TT. Methodology: MMu MMo JiN MC KI TT. Project administration: TT. Resources: TT JiN KI. Supervision: TT. Validation: MMu MMo CT MC KD KI. Visualization: MMu TT. Writing – original draft: MMu TT. Writing – review & editing: MMu TT MC JiN KI.