About the Authors

Qiongshi Lu

Contributed equally to this work with: Qiongshi Lu, Ryan Lee Powles

Affiliation Department of Biostatistics, Yale School of Public Health, New Haven, Connecticut, United States of America

Ryan Lee Powles

Contributed equally to this work with: Qiongshi Lu, Ryan Lee Powles

Affiliation Program of Computational Biology and Bioinformatics, Yale University, New Haven, Connecticut, United States of America

Qian Wang

Affiliation Program of Computational Biology and Bioinformatics, Yale University, New Haven, Connecticut, United States of America

Beixin Julie He

Affiliation Division of Cardiovascular Medicine, Department of Internal Medicine, Yale School of Medicine, New Haven, Connecticut, United States of America

Hongyu Zhao

hongyu.zhao@yale.edu

Affiliations Department of Biostatistics, Yale School of Public Health, New Haven, Connecticut, United States of America, Program of Computational Biology and Bioinformatics, Yale University, New Haven, Connecticut, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: QL RLP HZ. Analyzed the data: QL RLP QW BJH. Wrote the paper: QL RLP HZ. Advised on statistical and genetic issues: HZ.