About the Authors

Ryutaro Uchi

Contributed equally to this work with: Ryutaro Uchi, Yusuke Takahashi

Affiliations Department of Surgery, Kyushu University Beppu Hospital, Beppu, Japan, Department of Otorhinolaryngology, Graduate School of Medical Sciences, Kyushu University, Fukuoka, Japan

Yusuke Takahashi

Contributed equally to this work with: Ryutaro Uchi, Yusuke Takahashi

Affiliations Department of Surgery, Kyushu University Beppu Hospital, Beppu, Japan, Department of Gastroenterological Surgery, Graduate School of Medicine, Osaka University, Suita, Japan

Atsushi Niida

aniida@ims.u-tokyo.ac.jp (AN); kmimori@beppu.kyushu-u.ac.jp (KM)

‡ AN and KM also contributed equally to this work.

Affiliation Laboratory of DNA Information Analysis, Human Genome Center, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Teppei Shimamura

Affiliation Laboratory of DNA Information Analysis, Human Genome Center, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Hidenari Hirata

Affiliation Department of Surgery, Kyushu University Beppu Hospital, Beppu, Japan

Keishi Sugimachi

Affiliation Department of Surgery, Kyushu University Beppu Hospital, Beppu, Japan

Genta Sawada

Affiliations Department of Surgery, Kyushu University Beppu Hospital, Beppu, Japan, Department of Gastroenterological Surgery, Graduate School of Medicine, Osaka University, Suita, Japan

Takeshi Iwaya

Affiliation Department of Surgery, Iwate Medical University, Morioka, Japan

Junji Kurashige

Affiliation Department of Surgery, Kyushu University Beppu Hospital, Beppu, Japan

Yoshiaki Shinden

Affiliation Department of Surgery, Kyushu University Beppu Hospital, Beppu, Japan

Tomohiro Iguchi

Affiliation Department of Surgery, Kyushu University Beppu Hospital, Beppu, Japan

Hidetoshi Eguchi

Affiliation Department of Surgery, Kyushu University Beppu Hospital, Beppu, Japan

Kenichi Chiba

Affiliation Laboratory of DNA Information Analysis, Human Genome Center, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Yuichi Shiraishi

Affiliation Laboratory of DNA Information Analysis, Human Genome Center, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Genta Nagae

Affiliation Genome Science Laboratory, Research Center for Advanced Science and Technology, University of Tokyo, Tokyo, Japan

Kenichi Yoshida

Affiliation Department of Biology, Faculty of Sciences, Kyushu University, Fukuoka, Japan

Yasunobu Nagata

Affiliation Department of Biology, Faculty of Sciences, Kyushu University, Fukuoka, Japan

Hiroshi Haeno

Affiliation Department of Pathology and Tumor Biology, Kyoto University, Kyoto, Japan

Hirofumi Yamamoto

Affiliation Department of Gastroenterological Surgery, Graduate School of Medicine, Osaka University, Suita, Japan

Hideshi Ishii

Affiliation Department of Gastroenterological Surgery, Graduate School of Medicine, Osaka University, Suita, Japan

Yuichiro Doki

Affiliation Department of Gastroenterological Surgery, Graduate School of Medicine, Osaka University, Suita, Japan

Hisae Iinuma

Affiliation Department of Surgery, Teikyo University School of Medicine, Tokyo, Japan

Shin Sasaki

Affiliation Department of Surgery, Omori Red Cross Hospital, Tokyo, Japan

Satoshi Nagayama

Affiliation Gastroenterological Center, Department of Gastroenterological Surgery, Cancer Institute Hospital, Japanese Foundation for Cancer Research, Tokyo, Japan

Kazutaka Yamada

Affiliation Department of Surgery, Takano Hospital, Kumamoto, Japan

Shinichi Yachida

Affiliation Division of Cancer Genomics, National Cancer Center Research Institute, Tokyo, Japan

Mamoru Kato

Affiliation Division of Cancer Genomics, National Cancer Center Research Institute, Tokyo, Japan

Tatsuhiro Shibata

Affiliation Division of Cancer Genomics, National Cancer Center Research Institute, Tokyo, Japan

Eiji Oki

Affiliation Department of Surgery and Science, Graduate School of Medical Sciences, Kyushu University, Fukuoka, Japan

Hiroshi Saeki

Affiliation Department of Surgery and Science, Graduate School of Medical Sciences, Kyushu University, Fukuoka, Japan

Ken Shirabe

Affiliation Department of Surgery and Science, Graduate School of Medical Sciences, Kyushu University, Fukuoka, Japan

Yoshinao Oda

Affiliation Department of Anatomical Pathology, Graduate School of Medical Sciences, Kyushu University, Fukuoka, Japan

Yoshihiko Maehara

Affiliation Department of Surgery and Science, Graduate School of Medical Sciences, Kyushu University, Fukuoka, Japan

Shizuo Komune

Affiliation Department of Otorhinolaryngology, Graduate School of Medical Sciences, Kyushu University, Fukuoka, Japan

Masaki Mori

Affiliation Department of Gastroenterological Surgery, Graduate School of Medicine, Osaka University, Suita, Japan

Yutaka Suzuki

Affiliation Medical Genome Sciences, Graduate School of Frontier Sciences, University of Tokyo, Kashiwa-shi, Chiba, Japan

Ken Yamamoto

Affiliation Department of Medical Chemistry, Kurume University School of Medicine, Kurume, Japan

Hiroyuki Aburatani

Affiliation Genome Science Laboratory, Research Center for Advanced Science and Technology, University of Tokyo, Tokyo, Japan

Seishi Ogawa

Affiliation Department of Biology, Faculty of Sciences, Kyushu University, Fukuoka, Japan

Satoru Miyano

Affiliation Laboratory of DNA Information Analysis, Human Genome Center, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Koshi Mimori

aniida@ims.u-tokyo.ac.jp (AN); kmimori@beppu.kyushu-u.ac.jp (KM)

‡ AN and KM also contributed equally to this work.

Affiliation Department of Surgery, Kyushu University Beppu Hospital, Beppu, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YT AN KM MM SM TS MK SY KYamad SN SS HIi. Performed the experiments: YT RU GS YSu KYo HA GN KYamam YN TIw JK. Analyzed the data: RU AN HHi TS GS HHa. Contributed reagents/materials/analysis tools: AN KC YShin KSu HE TIg YShir MM SK YD HY YM EO YO KSh HS SO HIs. Wrote the paper: YT RU AN SO.