About the Authors

Rebecca K. Holmes

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Alex C. Tuck

Affiliations FMI Basel, Basel, Switzerland, European Molecular Biology Laboratory, European Bioinformatics Institute (EMBL-EBI), Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Chenchen Zhu

Affiliation EMBL, Heidelberg, Germany

Hywel R. Dunn-Davies

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Grzegorz Kudla

Affiliation The Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital, Edinburgh, Scotland, United Kingdom

Sandra Clauder-Munster

Affiliation EMBL, Heidelberg, Germany

Sander Granneman

Affiliation SynthSys, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Lars M. Steinmetz

Affiliation EMBL, Heidelberg, Germany

Christine Guthrie

Affiliation Department of Biochemistry and Biophysics, University of California, San Francisco, San Francisco, California, United States of America

David Tollervey

d.tollervey@ed.ac.uk

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RKH CG DT. Performed the experiments: RKH CZ SCM. Analyzed the data: RH ACT HRDD GK SG LMS DT. Contributed reagents/materials/analysis tools: ACT GK SG. Wrote the paper: RKH ACT LMS GK CG DT.