About the Authors

Hanqian L. Carlson

Affiliation Skeletal Biology Program, Shriners Hospitals for Children, Portland, Oregon, United States of America

Jeffrey J. Quinn

Affiliation Program in Epithelial Biology, Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Yul W. Yang

Affiliation Program in Epithelial Biology, Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Chelsea K. Thornburg

Affiliation Department of Biochemistry & Molecular Biology, Michigan State University, East Lansing, Michigan, United States of America

Howard Y. Chang

Affiliation Program in Epithelial Biology, Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

H. Scott Stadler

hss@shcc.org

Affiliations Skeletal Biology Program, Shriners Hospitals for Children, Portland, Oregon, United States of America, Department of Molecular and Medical Genetics, Oregon Health & Science University, Portland, Oregon, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HSS HYC. Performed the experiments: HLC YWY CKT. Analyzed the data: HLC HSS JJQ. Contributed reagents/materials/analysis tools: HYC YWY. Wrote the paper: HSS HYC HLC.