About the Authors

Julian R. Homburger

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Andrés Moreno-Estrada

amoreno@langebio.cinvestav.mx (AME); Marta-Alarcon-Riquelme@omrf.org (MEAR); cdbustam@stanford.edu (CDB)

Affiliations Department of Genetics, Stanford University, Stanford, California, United States of America, Laboratorio Nacional de Genómica para la Biodiversidad (LANGEBIO), CINVESTAV, Irapuato, Guanajuato, Mexico

Christopher R. Gignoux

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Dominic Nelson

Affiliations McGill University and Genome Quebec Innovation Centre, Montreal, Quebec, Canada, Department of Human Genetics, McGill University, Montreal, Quebec, Canada

Elena Sanchez

Affiliation Arthritis and Clinical Immunology, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Patricia Ortiz-Tello

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Bernardo A. Pons-Estel

Affiliation Sanatorio Parque, Rosario, Argentina

Eduardo Acevedo-Vasquez

Affiliation Facultad de Medicina, Universidad Nacional Mayor de San Marcos, Hospital Nacional Guillermo Almenara Irigoyen, Lima, Peru

Pedro Miranda

Affiliation Centro de Estudios Reumatologicos, Santiago, Chile

Carl D. Langefeld

Affiliation Center for Public Health Genomics, Wake Forest School of Medicine, Winston-Salem, North Carolina, United States of America

Simon Gravel

Affiliations McGill University and Genome Quebec Innovation Centre, Montreal, Quebec, Canada, Department of Human Genetics, McGill University, Montreal, Quebec, Canada

Marta E. Alarcón-Riquelme

amoreno@langebio.cinvestav.mx (AME); Marta-Alarcon-Riquelme@omrf.org (MEAR); cdbustam@stanford.edu (CDB)

Affiliations Arthritis and Clinical Immunology, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America, GENYO, Centre for Genomics and Oncological Research: Pfizer/ University of Granada/ Andalusian Regional Government, Granada, Spain

Carlos D. Bustamante

amoreno@langebio.cinvestav.mx (AME); Marta-Alarcon-Riquelme@omrf.org (MEAR); cdbustam@stanford.edu (CDB)

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Competing Interests

I have read the journal's policy and the authors of this manuscript have the following competing interests: CDB is on the Scientific Advisory Boards of Ancestry.com, Personalis, Liberty Biosecurity, 23andMe’s ‘‘Roots into the Future’’ project and Etalon DX. He is also a founder and Chair of the SAB of IdentifyGenomics. None of these entities played a role in the design, interpretation, or presentation of these results.

Author Contributions

Conceived and designed the experiments: AME CDB MEAR JRH CRG. Performed the experiments: MEAR ES CDL. Analyzed the data: JRH CRG ES AME POT DN SG. Contributed reagents/materials/analysis tools: MEAR BAPE EAV PM CDL CDB. Wrote the paper: JRH AME CRG CDB.