About the Authors

Jingqun Ao

Contributed equally to this work with: Jingqun Ao, Yinnan Mu, Li-Xin Xiang, DingDing Fan, MingJi Feng, Shicui Zhang

Affiliation Key Laboratory of Marine Biogenetics and Resources, Third Institute of Oceanography, State Oceanic Administration, Fujian Collaborative Innovation Center for Exploitation and Utilization of Marine Biological Resources, Key Laboratory of Marine Genetic Resources of Fujian Province, Xiamen, P. R. China

Yinnan Mu

Contributed equally to this work with: Jingqun Ao, Yinnan Mu, Li-Xin Xiang, DingDing Fan, MingJi Feng, Shicui Zhang

Affiliation Key Laboratory of Marine Biogenetics and Resources, Third Institute of Oceanography, State Oceanic Administration, Fujian Collaborative Innovation Center for Exploitation and Utilization of Marine Biological Resources, Key Laboratory of Marine Genetic Resources of Fujian Province, Xiamen, P. R. China

Li-Xin Xiang

Contributed equally to this work with: Jingqun Ao, Yinnan Mu, Li-Xin Xiang, DingDing Fan, MingJi Feng, Shicui Zhang

Affiliation College of Life Sciences, ZheJiang University, Key Laboratory for Cell and Gene Engineering of Zhejiang Province, Hangzhou, ZheJiang, P. R. China

DingDing Fan

Contributed equally to this work with: Jingqun Ao, Yinnan Mu, Li-Xin Xiang, DingDing Fan, MingJi Feng, Shicui Zhang

Affiliation BGI-Tech, BGI-Shenzhen, Shenzhen, Guangdong, P. R. China

MingJi Feng

Contributed equally to this work with: Jingqun Ao, Yinnan Mu, Li-Xin Xiang, DingDing Fan, MingJi Feng, Shicui Zhang

Affiliation BGI-Tech, BGI-Shenzhen, Shenzhen, Guangdong, P. R. China

Shicui Zhang

Contributed equally to this work with: Jingqun Ao, Yinnan Mu, Li-Xin Xiang, DingDing Fan, MingJi Feng, Shicui Zhang

Affiliation Ocean University of China, Qingdao, Shandong, P. R. China

Qiong Shi

Affiliation BGI-Tech, BGI-Shenzhen, Shenzhen, Guangdong, P. R. China

Lv-Yun Zhu

Affiliation College of Life Sciences, ZheJiang University, Key Laboratory for Cell and Gene Engineering of Zhejiang Province, Hangzhou, ZheJiang, P. R. China

Ting Li

Affiliation Key Laboratory of Marine Biogenetics and Resources, Third Institute of Oceanography, State Oceanic Administration, Fujian Collaborative Innovation Center for Exploitation and Utilization of Marine Biological Resources, Key Laboratory of Marine Genetic Resources of Fujian Province, Xiamen, P. R. China

Yang Ding

Affiliation Key Laboratory of Marine Biogenetics and Resources, Third Institute of Oceanography, State Oceanic Administration, Fujian Collaborative Innovation Center for Exploitation and Utilization of Marine Biological Resources, Key Laboratory of Marine Genetic Resources of Fujian Province, Xiamen, P. R. China

Li Nie

Affiliation College of Life Sciences, ZheJiang University, Key Laboratory for Cell and Gene Engineering of Zhejiang Province, Hangzhou, ZheJiang, P. R. China

Qiuhua Li

Affiliation Key Laboratory of Marine Biogenetics and Resources, Third Institute of Oceanography, State Oceanic Administration, Fujian Collaborative Innovation Center for Exploitation and Utilization of Marine Biological Resources, Key Laboratory of Marine Genetic Resources of Fujian Province, Xiamen, P. R. China

Wei-ren Dong

Affiliation College of Life Sciences, ZheJiang University, Key Laboratory for Cell and Gene Engineering of Zhejiang Province, Hangzhou, ZheJiang, P. R. China

Liang Jiang

Affiliation College of Life Sciences, Shenzhen University, Shenzhen, Guangdong, P. R. China

Bing Sun

Affiliation Ocean University of China, Qingdao, Shandong, P. R. China

XinHui Zhang

Affiliation BGI-Tech, BGI-Shenzhen, Shenzhen, Guangdong, P. R. China

Mingyu Li

Affiliation Key Laboratory of Marine Biogenetics and Resources, Third Institute of Oceanography, State Oceanic Administration, Fujian Collaborative Innovation Center for Exploitation and Utilization of Marine Biological Resources, Key Laboratory of Marine Genetic Resources of Fujian Province, Xiamen, P. R. China

Hai-Qi Zhang

Affiliation College of Life Sciences, ZheJiang University, Key Laboratory for Cell and Gene Engineering of Zhejiang Province, Hangzhou, ZheJiang, P. R. China

ShangBo Xie

Affiliation BGI-Tech, BGI-Shenzhen, Shenzhen, Guangdong, P. R. China

YaBing Zhu

Affiliation BGI-Tech, BGI-Shenzhen, Shenzhen, Guangdong, P. R. China

XuanTing Jiang

Affiliation BGI-Tech, BGI-Shenzhen, Shenzhen, Guangdong, P. R. China

Xianhui Wang

Affiliation Key Laboratory of Marine Biogenetics and Resources, Third Institute of Oceanography, State Oceanic Administration, Fujian Collaborative Innovation Center for Exploitation and Utilization of Marine Biological Resources, Key Laboratory of Marine Genetic Resources of Fujian Province, Xiamen, P. R. China

Pengfei Mu

Affiliation Key Laboratory of Marine Biogenetics and Resources, Third Institute of Oceanography, State Oceanic Administration, Fujian Collaborative Innovation Center for Exploitation and Utilization of Marine Biological Resources, Key Laboratory of Marine Genetic Resources of Fujian Province, Xiamen, P. R. China

Wei Chen

Affiliation Key Laboratory of Marine Biogenetics and Resources, Third Institute of Oceanography, State Oceanic Administration, Fujian Collaborative Innovation Center for Exploitation and Utilization of Marine Biological Resources, Key Laboratory of Marine Genetic Resources of Fujian Province, Xiamen, P. R. China

Zhen Yue

Affiliation BGI-Tech, BGI-Shenzhen, Shenzhen, Guangdong, P. R. China

Zhuo Wang

Affiliation BGI-Tech, BGI-Shenzhen, Shenzhen, Guangdong, P. R. China

Jun Wang

wangj@genomics.org.cn (JW); shaojz@zju.edu.cn (JS); chenxinhua@tio.org.cn (XC)

Affiliation BGI-Tech, BGI-Shenzhen, Shenzhen, Guangdong, P. R. China

Jian-Zhong Shao

wangj@genomics.org.cn (JW); shaojz@zju.edu.cn (JS); chenxinhua@tio.org.cn (XC)

Affiliation College of Life Sciences, ZheJiang University, Key Laboratory for Cell and Gene Engineering of Zhejiang Province, Hangzhou, ZheJiang, P. R. China

Xinhua Chen

wangj@genomics.org.cn (JW); shaojz@zju.edu.cn (JS); chenxinhua@tio.org.cn (XC)

Affiliation Key Laboratory of Marine Biogenetics and Resources, Third Institute of Oceanography, State Oceanic Administration, Fujian Collaborative Innovation Center for Exploitation and Utilization of Marine Biological Resources, Key Laboratory of Marine Genetic Resources of Fujian Province, Xiamen, P. R. China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JA SZ QS JW JZS XC. Performed the experiments: YM YD QL JA TL ML XW PM WC. Analyzed the data: DF MF LXX LJ XZ SX YZ XJ ZY ZW YM JA QS SZ LYZ TL LN WrD BS HQZ. Wrote the paper: JA YM LXX DF SZ QS LYZ JZS XC.