About the Authors

Norihito Uemura

Affiliation Department of Neurology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Masato Koike

Affiliation Department of Cell Biology and Neuroscience, Juntendo University Graduate School of Medicine, Tokyo, Japan

Satoshi Ansai

Affiliation Division of Applied Biosciences, Kyoto University Graduate School of Agriculture, Kyoto, Japan

Masato Kinoshita

Affiliation Division of Applied Biosciences, Kyoto University Graduate School of Agriculture, Kyoto, Japan

Tomoko Ishikawa-Fujiwara

Affiliation Department of Radiation Biology and Medical Genetics, Osaka University Graduate School of Medicine, Suita, Japan

Hideaki Matsui

Current Address: Department of Neuroscience, Section of Integrative Physiology, Faculty of Medicine, University of Miyazaki, Miyazaki, Japan

Affiliation Department of Neurology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Kiyoshi Naruse

Affiliation National Institute for Basic Biology, Laboratory of Bioresources, Okazaki, Japan

Naoaki Sakamoto

Affiliation Department of Mathematical and Life Sciences, Hiroshima University Graduate School of Science, Higashi-Hiroshima, Japan

Yasuo Uchiyama

Affiliation Department of Cell Biology and Neuroscience, Juntendo University Graduate School of Medicine, Tokyo, Japan

Takeshi Todo

Affiliation Department of Radiation Biology and Medical Genetics, Osaka University Graduate School of Medicine, Suita, Japan

Shunichi Takeda

Affiliations Department of Radiation Genetics, Kyoto University Graduate School of Medicine, Kyoto, Japan, Core Research for Evolutional Science and Technology, Japan Science and Technology Agency, Kawaguchi, Japan

Hodaka Yamakado

‡ These authors are joint senior authors on this work.

Affiliations Department of Neurology, Kyoto University Graduate School of Medicine, Kyoto, Japan, Core Research for Evolutional Science and Technology, Japan Science and Technology Agency, Kawaguchi, Japan

Ryosuke Takahashi

ryosuket@kuhp.kyoto-u.ac.jp

‡ These authors are joint senior authors on this work.

Affiliations Department of Neurology, Kyoto University Graduate School of Medicine, Kyoto, Japan, Core Research for Evolutional Science and Technology, Japan Science and Technology Agency, Kawaguchi, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NU HM HY ST RT. Performed the experiments: NU MKo SA MKi TIF TT. Analyzed the data: NU MKo YU. Contributed reagents/materials/analysis tools: KN NS. Wrote the paper: NU MKo MKi HM HY RT.