About the Authors

Alicia R. Martin

Contributed equally to this work with: Alicia R. Martin, Helio A. Costa

Affiliation Stanford University School of Medicine, Department of Genetics, Stanford, California, United States of America

Helio A. Costa

Contributed equally to this work with: Alicia R. Martin, Helio A. Costa

Affiliation Stanford University School of Medicine, Department of Genetics, Stanford, California, United States of America

Tuuli Lappalainen

Affiliation Stanford University School of Medicine, Department of Genetics, Stanford, California, United States of America

Brenna M. Henn

Affiliation Stony Brook University, SUNY, Department of Ecology and Evolution, Stony Brook, New York, United States of America

Jeffrey M. Kidd

Affiliation University of Michigan School of Medicine, Department of Human Genetics, Ann Arbor, Michigan, United States of America

Muh-Ching Yee

Affiliation Stanford University School of Medicine, Department of Genetics, Stanford, California, United States of America

Fabian Grubert

Affiliation Stanford University School of Medicine, Department of Genetics, Stanford, California, United States of America

Howard M. Cann

Affiliation Foundation Jean Dausset, Centre d'Etude du Polymorphisme Humain, Paris, France

Michael Snyder

Affiliation Stanford University School of Medicine, Department of Genetics, Stanford, California, United States of America

Stephen B. Montgomery

Affiliations Stanford University School of Medicine, Department of Genetics, Stanford, California, United States of America, Stanford University School of Medicine, Department of Pathology, Stanford, California, United States of America

Carlos D. Bustamante

cdbustam@stanford.edu

Affiliation Stanford University School of Medicine, Department of Genetics, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ARM HAC MCY MS CDB TL SBM. Performed the experiments: MCY FG. Analyzed the data: ARM HAC. Contributed reagents/materials/analysis tools: ARM HAC CDB MS SBM JMK HMC BMH. Wrote the paper: ARM HAC. Contributed to final draft: ARM HAC TL SBM BMH JMK CDB.