About the Authors

Wayne N. Frankel

wayne.frankel@jax.org

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Connie L. Mahaffey

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Tracy C. McGarr

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Barbara J. Beyer

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Verity A. Letts

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WNF. Performed the experiments: WNF CLM TCM BJB VAL. Analyzed the data: WNF CLM. Wrote the paper: WNF CLM.