About the Authors

Jason A. Hodgson

Affiliation Department of Life Sciences, Silwood Park Campus, Imperial College London, Ascot, Berkshire, United Kingdom

Connie J. Mulligan

Affiliation Department of Anthropology and the Genetics Institute, University of Florida, Gainesville, Florida, United States of America

Ali Al-Meeri

Affiliation Department of Biochemistry and Molecular Biology, Sana'a University, Sana'a, Yemen

Ryan L. Raaum

ryan.raaum@lehman.cuny.edu

Affiliations Department of Anthropology, Lehman College and The Graduate Center, The City University of New York, Bronx, New York, New York, United States of America, The New York Consortium in Evolutionary Primatology (NYCEP), New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JAH RLR. Performed the experiments: JAH CJM RLR. Analyzed the data: JAH RLR. Contributed reagents/materials/analysis tools: CJM AAM. Wrote the paper: JAH RLR.