About the Authors

Zhe Zhang

Contributed equally to this work with: Zhe Zhang, Chan-Juan Hao

Affiliations State Key Laboratory of Molecular Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, China, Graduate School of Chinese Academy of Sciences, Beijing, China

Chan-Juan Hao

Contributed equally to this work with: Zhe Zhang, Chan-Juan Hao

Affiliations State Key Laboratory of Molecular Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, China, Graduate School of Chinese Academy of Sciences, Beijing, China, Reproductive and Genetic Center, National Research Institute for Family Planning, Beijing, China

Chang-Gui Li

Affiliation The Affiliated Hospital of Qingdao University Medical College, Qingdao, China

Dong-Jie Zang

Affiliation State Key Laboratory of Molecular Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, China

Jing Zhao

Affiliation Model Animal Research Center, Nanjing University, Nanjing, China

Xiao-Nan Li

Affiliation Nanjing Children's Hospital, Nanjing Medical University, Nanjing, China

Ai-Hua Wei

Affiliation Beijing Tongren Hospital, Capital Medical University, Beijing, China

Zong-Bo Wei

Affiliations State Key Laboratory of Molecular Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, China, Graduate School of Chinese Academy of Sciences, Beijing, China

Lin Yang

Affiliation State Key Laboratory of Molecular Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, China

Xin He

Affiliation State Key Laboratory of Molecular Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, China

Xue-Chu Zhen

Affiliation Department of Pharmacology, Soochow University College of Pharmaceutical Sciences, Suzhou, China

Xiang Gao

Affiliation Model Animal Research Center, Nanjing University, Nanjing, China

John R. Speakman

Affiliations State Key Laboratory of Molecular Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, China, Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen, Scotland, United Kingdom

Wei Li

wli@genetics.ac.cn

Affiliation State Key Laboratory of Molecular Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WL JRS XG XCZ CGL XNL. Performed the experiments: ZZ CJH DJZ AHW ZBW JZ XH LY. Analyzed the data: WL JRS XG XCZ CGL XNL. Contributed reagents/materials/analysis tools: ZBW JRS. Wrote the paper: WL ZZ CJH JRS.