About the Authors

Xu Wang

Affiliations Department of Molecular Biology and Genetics, Cornell University, Ithaca, New York, United States of America, Cornell Center for Comparative and Population Genomics, Cornell University, Ithaca, New York, United States of America

David Wheeler

Current address: Institute of Fundamental Sciences, Massey University, Palmerston North, New Zealand.

Affiliation Department of Biology, University of Rochester, Rochester, New York, United States of America

Amanda Avery

Affiliation Department of Biology, University of Rochester, Rochester, New York, United States of America

Alfredo Rago

Affiliation School of Biosciences, The University of Birmingham, Birmingham, United Kingdom

Jeong-Hyeon Choi

Affiliation Cancer Center, Department of Biostatistics and Epidemiology, Georgia Regents University, Augusta, Georgia, United States of America

John K. Colbourne

Affiliation School of Biosciences, The University of Birmingham, Birmingham, United Kingdom

Andrew G. Clark

ac347@cornell.edu (AGC); werr@mail.rochester.edu (JHW)

Affiliations Department of Molecular Biology and Genetics, Cornell University, Ithaca, New York, United States of America, Cornell Center for Comparative and Population Genomics, Cornell University, Ithaca, New York, United States of America

John H. Werren

ac347@cornell.edu (AGC); werr@mail.rochester.edu (JHW)

Affiliation Department of Biology, University of Rochester, Rochester, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XW JKC AGC JHW. Performed the experiments: XW AA. Analyzed the data: XW DW AR JHC JKC AGC JHW. Contributed reagents/materials/analysis tools: JKC AGC JHW. Wrote the paper: XW JKC AGC JHW.