About the Authors

Girish Deshpande

Affiliation Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Keren Zhou

Affiliation Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Joy Y. Wan

Affiliation Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Jana Friedrich

Affiliation Centre for Organismal Studies (COS) University of Heidelberg, Heidelberg, Germany

Nicholas Jourjine

Affiliation Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Daniel Smith

Affiliation Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Paul Schedl

pschedl@princeton.edu

Affiliations Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America, Institute of Gene Biology RAS, Moscow, Russia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GD PS. Performed the experiments: GD KZ JYW JF NJ DS. Analyzed the data: GD KZ JYW JF NJ DS. Contributed reagents/materials/analysis tools: GD KZ JYW JF NJ DS. Wrote the paper: GD DS PS.