About the Authors

Helen Strutt

Affiliation MRC Centre for Developmental and Biomedical Genetics and Department of Biomedical Science, University of Sheffield, Western Bank, Sheffield, United Kingdom

Vickie Thomas-MacArthur

Affiliation MRC Centre for Developmental and Biomedical Genetics and Department of Biomedical Science, University of Sheffield, Western Bank, Sheffield, United Kingdom

David Strutt

d.strutt@sheffield.ac.uk

Affiliation MRC Centre for Developmental and Biomedical Genetics and Department of Biomedical Science, University of Sheffield, Western Bank, Sheffield, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HS DS. Performed the experiments: HS VTM DS. Analyzed the data: HS DS. Wrote the paper: HS DS.