About the Authors

Harinder Manku

Affiliation Department of Medical & Molecular Genetics, King's College London School of Medicine, Guy's Hospital, London, United Kingdom

Carl D. Langefeld

Affiliation Wake Forest School of Medicine, Winston-Salem, North Carolina, United States of America

Sandra G. Guerra

Affiliation Centre for Rheumatology & Connective Tissue Diseases, Royal Free & University College Medical School, London, United Kingdom

Talat H. Malik

Affiliation Division of Immunology and Inflammation, Imperial College, London, United Kingdom

Marta Alarcon-Riquelme

Affiliation Centro Pfizer-Universidad de Granada-Junta de Andalucía de Genómica e Investigaciones Oncológicas, Granada, Spain

Juan-Manuel Anaya

Affiliation Center for Autoimmune Diseases Research, Universidad del Rosario, Bogota, Colombia

Sang-Cheol Bae

Affiliation Hospital for Rheumatic Diseases, Hanyang University, Seoul, South Korea

Susan A. Boackle

Affiliation Division of Rheumatology, University of Colorado Denver, Aurora, Colorado, United States of America

Elizabeth E. Brown

Affiliation Department of Epidemiology, University of Alabama at Birmingham, Birmingham, Alabama, United States of America

Lindsey A. Criswell

Affiliation Rosalind Russell Medical Research Center for Arthritis, University of California San Francisco, San Francisco, California, United States of America

Barry I. Freedman

Affiliation Department of Internal Medicine, Wake Forest School of Medicine, Winston-Salem, North Carolina, United States of America

Patrick M. Gaffney

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Peter A. Gregersen

Affiliation The Robert S. Boas Center for Genomics and Human Genetics, Feinstein Institute for Medical Research, North Shore LIJ Health System, Manhasset, New York, United States of America

Joel M. Guthridge

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Sang-Hoon Han

Affiliation Hospital for Rheumatic Diseases, Hanyang University, Seoul, South Korea

John B. Harley

Affiliation Division of Rheumatology, Cincinnati Children's Hospital Medical Centre, Cincinnati, Ohio, United States of America

Chaim O. Jacob

Affiliation The Lupus Genetics Group, Department of Medicine, University of Southern California, Los Angeles, California, United States of America

Judith A. James

Affiliations Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America, Department of Medicine, University of Oklahoma Healthy Sciences Center, Oklahoma City, Oklahoma, United States of America

Diane L. Kamen

Affiliation Division of Rheumatology, Medical University of South Carolina, Charleston, South Carolina, United States of America

Kenneth M. Kaufman

Affiliation Division of Rheumatology, Cincinnati Children's Hospital Medical Centre, Cincinnati, Ohio, United States of America

Jennifer A. Kelly

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Javier Martin

Affiliation Instituto de Parasitologia y Biomedicina Lopez-Neyra, Consejo Superior de Investigaciones Cientificas, Granada, Spain

Joan T. Merrill

Affiliation Clinical Pharmacology, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Kathy L. Moser

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Timothy B. Niewold

Affiliation Divisions of Rheumatology and Immunology, Mayo Clinic, Rochester, Minnesota, United States of America

So-Yeon Park

Affiliation Hospital for Rheumatic Diseases, Hanyang University, Seoul, South Korea

Bernardo A. Pons-Estel

Affiliation Sanatorio Parque, Rosario, Argentina

Amr H. Sawalha

Affiliation Division of Rheumatology, Department of Internal Medicine, University of Michigan, Ann Arbor, Michigan, United States of America

R. Hal Scofield

Affiliations Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America, Department of Medicine, University of Oklahoma Healthy Sciences Center, Oklahoma City, Oklahoma, United States of America

Nan Shen

Affiliation Shanghai Institute for Biological Sciences, Chinese Academy of Sciences, Shanghai, China

Anne M. Stevens

Affiliation Center for Immunity and Immunotherapies, Seattle Children's Research Institute, Seattle, Washington, United States of America

Celi Sun

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Gary S. Gilkeson

Affiliation Division of Rheumatology and Immunology, Medical University of South Carolina, Charleston, South Carolina, United States of America

Jeff C. Edberg

Affiliation Division of Rheumatology and Immunology, Medical University of South Carolina, Charleston, South Carolina, United States of America

Robert P. Kimberly

Affiliation Division of Clinical Immunology and Rheumatology, Department of Medicine, University of Alabama at Birmingham, Birmingham, Alabama, United States of America

Swapan K. Nath

Affiliation Arthritis and Clinical Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Betty P. Tsao

Affiliation Division of Rheumatology, Department of Medicine, David Geffen School of Medicine at UCLA, Los Angeles, California, United States of America

Tim J. Vyse

tim.vyse@kcl.ac.uk

Affiliation Department of Medical & Molecular Genetics, King's College London School of Medicine, Guy's Hospital, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HM TJV CDL JBH BPT SKN. Performed the experiments: HM JAK KMK SGG THM. Analyzed the data: HM CDL. Contributed reagents/materials/analysis tools: TJV CDL JBH MAR JMA SCB SAB EEB LAC BIF PMG PAG JMG SHH COJ JAJ DLK KMK JAK JM JTM KLM TBN SYP BAPE AHS RHS NS AMS CS GSG JCE RPK BPT. Wrote the paper: HM.