About the Authors

Noah Zaitlen

noah.zaitlen@ucsf.edu (NZ); aprice@hsph.harvard.edu (ALP)

Affiliation Department of Medicine, Lung Biology Center, University of California San Francisco, San Francisco, California, United States of America

Peter Kraft

Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America, Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America

Nick Patterson

Affiliation Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America

Bogdan Pasaniuc

Affiliation Interdepartmental Program in Bioinformatics Pathology and Laboratory Medicine, University of California Los Angeles, Los Angeles, California, United States of America

Gaurav Bhatia

Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America, Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America

Samuela Pollack

Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America, Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America

Alkes L. Price

noah.zaitlen@ucsf.edu (NZ); aprice@hsph.harvard.edu (ALP)

Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America, Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NZ NP ALP. Performed the experiments: NZ SP. Analyzed the data: NZ PK NP BP GB ALP. Wrote the paper: NZ ALP.