About the Authors

Allison C. Billi

Contributed equally to this work with: Allison C. Billi, Mallory A. Freeberg

Affiliations Life Sciences Institute, University of Michigan, Ann Arbor, Michigan, United States of America, Department of Human Genetics, University of Michigan, Ann Arbor, Michigan, United States of America

Mallory A. Freeberg

Contributed equally to this work with: Allison C. Billi, Mallory A. Freeberg

Affiliations Life Sciences Institute, University of Michigan, Ann Arbor, Michigan, United States of America, Department of Computational Medicine and Bioinformatics, University of Michigan, Ann Arbor, Michigan, United States of America

Amanda M. Day

Affiliations Life Sciences Institute, University of Michigan, Ann Arbor, Michigan, United States of America, Department of Cellular and Molecular Biology, University of Michigan, Ann Arbor, Michigan, United States of America

Sang Young Chun

Affiliations Life Sciences Institute, University of Michigan, Ann Arbor, Michigan, United States of America, Department of Computational Medicine and Bioinformatics, University of Michigan, Ann Arbor, Michigan, United States of America

Vishal Khivansara

Affiliation Life Sciences Institute, University of Michigan, Ann Arbor, Michigan, United States of America

John K. Kim

jnkim@umich.edu

Affiliations Life Sciences Institute, University of Michigan, Ann Arbor, Michigan, United States of America, Department of Human Genetics, University of Michigan, Ann Arbor, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MAF ACB JKK. Performed the experiments: ACB AMD VK. Analyzed the data: MAF SYC. Wrote the paper: ACB MAF JKK.