About the Authors

Bethan E. Phillips

Affiliation Graduate Entry Medicine and Health, Royal Derby Hospital, Derbyshire, United Kingdom

John P. Williams

Affiliation Graduate Entry Medicine and Health, Royal Derby Hospital, Derbyshire, United Kingdom

Thomas Gustafsson

Affiliation Karolinska Institutet, Stockholm, Sweden

Claude Bouchard

Affiliation Pennington Biomedical Research Center, Baton Rouge, Louisiana, United States of America

Tuomo Rankinen

Affiliation Pennington Biomedical Research Center, Baton Rouge, Louisiana, United States of America

Steen Knudsen

Affiliation Medical Prognosis Institute, Hørsholm, Denmark

Kenneth Smith

Affiliation Graduate Entry Medicine and Health, Royal Derby Hospital, Derbyshire, United Kingdom

James A. Timmons

Contributed equally to this work with: James A. Timmons, Philip J. Atherton

Jamie.Timmons@gmail.com

Affiliations XRgenomics Ltd., Leicestershire, United Kingdom, Loughborough University, Loughborough, United Kingdom

Philip J. Atherton

Contributed equally to this work with: James A. Timmons, Philip J. Atherton

Affiliation Graduate Entry Medicine and Health, Royal Derby Hospital, Derbyshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KS PJA JAT CB TG. Performed the experiments: BEP JPW CB KS JAT PJA. Analyzed the data: BEP KS JAT PJA. Contributed reagents/materials/analysis tools: CB TR TG. Wrote the paper: BEP TG CB SK JAT PJA.