About the Authors

Sandra Beleza

sbeleza@stanford.edu (SB); gbarsh@hudsonalpha.org (GSB)

Affiliations Department of Genetics, Stanford University School of Medicine, Stanford, California, United States of America, Instituto de Patologia e Imunologia Molecular da Universidade do Porto (IPATIMUP), Porto, Portugal

Nicholas A. Johnson

Affiliation Department of Statistics, Stanford University, Stanford, California, United States of America

Sophie I. Candille

Affiliation Department of Genetics, Stanford University School of Medicine, Stanford, California, United States of America

Devin M. Absher

Affiliation HudsonAlpha Institute for Biotechnology, Huntsville, Alabama, United States of America

Marc A. Coram

Affiliation Department of Health Research and Policy, Stanford University School of Medicine, Stanford, California, United States of America

Jailson Lopes

Affiliations Instituto de Patologia e Imunologia Molecular da Universidade do Porto (IPATIMUP), Porto, Portugal, Centro de Investigação em Biodiversidade e Recursos Genéticos (CIBIO), Vairão, Portugal, Universidade de Cabo Verde (Uni-CV), Praia, Santiago, Cabo Verde

Joana Campos

Affiliation Instituto de Patologia e Imunologia Molecular da Universidade do Porto (IPATIMUP), Porto, Portugal

Isabel Inês Araújo

Affiliation Universidade de Cabo Verde (Uni-CV), Praia, Santiago, Cabo Verde

Tovi M. Anderson

Affiliation Department of Genetics, Stanford University School of Medicine, Stanford, California, United States of America

Bjarni J. Vilhjálmsson

Affiliation Gregor Mendel Institute, Austrian Academy of Sciences, Vienna, Austria

Magnus Nordborg

Affiliation Gregor Mendel Institute, Austrian Academy of Sciences, Vienna, Austria

António Correia e Silva

Affiliation Universidade de Cabo Verde (Uni-CV), Praia, Santiago, Cabo Verde

Mark D. Shriver

Affiliation Department of Anthropology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Jorge Rocha

Affiliations Instituto de Patologia e Imunologia Molecular da Universidade do Porto (IPATIMUP), Porto, Portugal, Centro de Investigação em Biodiversidade e Recursos Genéticos (CIBIO), Vairão, Portugal, Departamento de Biologia, Faculdade de Ciências da Universidade do Porto, Porto, Portugal

Gregory S. Barsh

Contributed equally to this work with: Gregory S. Barsh, Hua Tang

sbeleza@stanford.edu (SB); gbarsh@hudsonalpha.org (GSB)

Affiliations Department of Genetics, Stanford University School of Medicine, Stanford, California, United States of America, HudsonAlpha Institute for Biotechnology, Huntsville, Alabama, United States of America

Hua Tang

Contributed equally to this work with: Gregory S. Barsh, Hua Tang

Affiliation Department of Genetics, Stanford University School of Medicine, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SB JR HT ACeS MDS GSB. Performed the experiments: SB JC JL IIA TMA JR DMA. Analyzed the data: SB NAJ SIC HT. Contributed reagents/materials/analysis tools: MAC NAJ BJV. Wrote the paper: SB HT GSB. Conceptualized and supported initiation of the study: ACeS MDS. Carried out the EMMAX analysis: BJV MN.