About the Authors

Zheng Hu

Affiliations Center for Computational Biology and Laboratory of Disease Genomics and Individualized Medicine, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China, Graduate University of Chinese Academy of Sciences, Beijing, China

Yun-Xin Fu

Affiliation Human Genetics Center and Division of Biostatistics, School of Public Health, University of Texas Health Science Center at Houston, Houston, Texas, United States of America

Anthony J. Greenberg

Affiliation Departments of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Chung-I Wu

weiweizhai@big.ac.cn (WZ); ciwu@uchicago.edu (C-IW)

Affiliations Center for Computational Biology and Laboratory of Disease Genomics and Individualized Medicine, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China, Department of Ecology and Evolution, University of Chicago, Chicago, Illinois, United States of America

Weiwei Zhai

weiweizhai@big.ac.cn (WZ); ciwu@uchicago.edu (C-IW)

Affiliations Center for Computational Biology and Laboratory of Disease Genomics and Individualized Medicine, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China, National Center for Mathematics and Interdisciplinary Sciences, Chinese Academy of Sciences, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WZ ZH. Performed the experiments: ZH Y-XF C-IW WZ. Analyzed the data: ZH Y-XF C-IW WZ. Wrote the paper: ZH Y-XF AJG C-IW WZ.