About the Authors

Yan Wang

Contributed equally to this work with: Yan Wang, Wen-yan Zhang

Affiliation State Key Laboratory of Microbial Technology, School of Life Science, Shandong University, Jinan, China

Wen-yan Zhang

Contributed equally to this work with: Yan Wang, Wen-yan Zhang

Affiliation State Key Laboratory of Microbial Technology, School of Life Science, Shandong University, Jinan, China

Zheng Zhang

Affiliation State Key Laboratory of Microbial Technology, School of Life Science, Shandong University, Jinan, China

Jian Li

Affiliation State Key Laboratory of Microbial Technology, School of Life Science, Shandong University, Jinan, China

Zhi-feng Li

Affiliation State Key Laboratory of Microbial Technology, School of Life Science, Shandong University, Jinan, China

Zai-gao Tan

Affiliation State Key Laboratory of Microbial Technology, School of Life Science, Shandong University, Jinan, China

Tian-tian Zhang

Affiliation State Key Laboratory of Microbial Technology, School of Life Science, Shandong University, Jinan, China

Zhi-hong Wu

Affiliation State Key Laboratory of Microbial Technology, School of Life Science, Shandong University, Jinan, China

Hong Liu

Affiliation State Key Laboratory of Microbial Technology, School of Life Science, Shandong University, Jinan, China

Yue-zhong Li

lilab@sdu.edu.cn

Affiliation State Key Laboratory of Microbial Technology, School of Life Science, Shandong University, Jinan, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YL YW WZ. Performed the experiments: YW WZ ZZ JL TZ ZT. Analyzed the data: YL YW WZ ZZ ZL ZW HL. Wrote the paper: YL YW ZZ WZ.