About the Authors

Adam Kiezun

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America, Cancer Program, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America

Sara L. Pulit

Affiliation Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America

Laurent C. Francioli

Affiliation Departments of Medical Genetics and of Epidemiology, University Medical Center Utrecht, Utrecht, The Netherlands

Freerk van Dijk

Affiliation Department of Genetics, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Morris Swertz

Affiliation Department of Genetics, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Dorret I. Boomsma

Affiliation Department of Biological Psychology, VU University, Amsterdam, The Netherlands

Cornelia M. van Duijn

Affiliation Department of Epidemiology, Erasmus Medical Center, Rotterdam, The Netherlands

P. Eline Slagboom

Affiliation Department of Medical Statistics and Bioinformatics, Leiden University Medical Center, Leiden, The Netherlands

G. J. B. van Ommen

Affiliation Department of Human Genetics, Leiden University Medical Center, Leiden, The Netherlands

Cisca Wijmenga

Affiliation Department of Genetics, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands

Genome of the Netherlands Consortium

Membership of the Genome of the Netherlands Consortium is provided in the Acknowledgments.

Paul I. W. de Bakker

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America, Departments of Medical Genetics and of Epidemiology, University Medical Center Utrecht, Utrecht, The Netherlands

Shamil R. Sunyaev

ssunyaev@rics.bwh.harvard.edu

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SRS AK SLP PIWdB. Performed the experiments: AK SLP LCF. Analyzed the data: SRS AK SLP PIWdB. Contributed reagents/materials/analysis tools: LCF FvD MS DIB CMvD PES GJBvO CW PIWdB Genome of the Netherlands Consortium. Wrote the paper: SRS AK PIWdB.