About the Authors

Gibran Hemani

g.hemani@uq.edu.au

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Science, University of Edinburgh, Edinburgh, United Kingdom

Sara Knott

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital, Edinburgh, United Kingdom

Chris Haley

Affiliations The Roslin Institute and Royal (Dick) School of Veterinary Science, University of Edinburgh, Edinburgh, United Kingdom, Institute for Evolutionary Biology, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GH SK CH. Performed the experiments: GH. Analyzed the data: GH. Wrote the paper: GH.