About the Authors

Kei Semba

Contributed equally to this work with: Kei Semba, Kimi Araki

Affiliations Division of Developmental Genetics, Institute of Resource Development and Analysis, Kumamoto University, Kumamoto, Japan, Department of Orthopedic and Neuro-Musculoskeletal Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Kimi Araki

Contributed equally to this work with: Kei Semba, Kimi Araki

Affiliation Division of Developmental Genetics, Institute of Resource Development and Analysis, Kumamoto University, Kumamoto, Japan

Ken-ichirou Matsumoto

Affiliations Division of Developmental Genetics, Institute of Resource Development and Analysis, Kumamoto University, Kumamoto, Japan, Department of Orthopedic and Neuro-Musculoskeletal Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Hiroko Suda

Affiliations Division of Developmental Genetics, Institute of Resource Development and Analysis, Kumamoto University, Kumamoto, Japan, Department of Transplantation/Pediatric Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Takashi Ando

Affiliations Division of Developmental Genetics, Institute of Resource Development and Analysis, Kumamoto University, Kumamoto, Japan, Department of Orthopedic and Neuro-Musculoskeletal Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Akira Sei

Affiliation Department of Orthopedic and Neuro-Musculoskeletal Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Hiroshi Mizuta

Affiliation Department of Orthopedic and Neuro-Musculoskeletal Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Katsumasa Takagi

Affiliation Department of Orthopedic and Neuro-Musculoskeletal Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Mai Nakahara

Affiliation Division of Developmental Genetics, Institute of Resource Development and Analysis, Kumamoto University, Kumamoto, Japan

Mayumi Muta

Affiliation Division of Developmental Genetics, Institute of Resource Development and Analysis, Kumamoto University, Kumamoto, Japan

Gen Yamada

Affiliation Department of Organ Formation, Institute of Resource Development and Analysis, Kumamoto University, Kumamoto, Japan

Naomi Nakagata

Affiliation Division of Reproductive Engineering, Institute of Resource Development and Analysis, Kumamoto University, Kumamoto, Japan

Aritoshi Iida

Affiliation Laboratory for Bone and Joint Diseases, RIKEN Center for Genomic Medicine, Tokyo, Japan

Shiro Ikegawa

Affiliation Laboratory for Bone and Joint Diseases, RIKEN Center for Genomic Medicine, Tokyo, Japan

Yusuke Nakamura

Affiliation Human Genome Center, Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Masatake Araki

Affiliation Division of Bioinformatics, Institute of Resource Development and Analysis, Kumamoto University, Kumamoto, Japan

Kuniya Abe

Affiliation Technology Development Team for Mammalian Cellular Dynamics, BioResource Center, RIKEN, Tsukuba, Ibaraki, Japan

Ken-ichi Yamamura

yamamura@gpo.kumamoto-u.ac.jp

Affiliation Division of Developmental Genetics, Institute of Resource Development and Analysis, Kumamoto University, Kumamoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: K Semba, K Araki, M Araki, K Abe, K Yamamura. Performed the experiments: K Semba, K Araki, K Matsumoto, H Suda, T Ando, M Nakahara, M Muta, A Iida, K Abe. Analyzed the data: K Semba, K Araki, A Sei, H Mizuta, K Takagi, G Yamada, A Iida, S Ikegawa, Y Nakamura, K Abe, K Yamamura. Contributed reagents/materials/analysis tools: K Semba, K Araki, N Nakagata, A Iida, Y Nakamura, K Abe. Wrote the paper: K Semba, K Yamamura.