About the Authors

Vinod Kumar

Affiliation Department of Genetics, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands

Harm-Jan Westra

Contributed equally to this work with: Harm-Jan Westra, Juha Karjalainen, Daria V. Zhernakova

Affiliation Department of Genetics, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands

Juha Karjalainen

Contributed equally to this work with: Harm-Jan Westra, Juha Karjalainen, Daria V. Zhernakova

Affiliation Department of Genetics, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands

Daria V. Zhernakova

Contributed equally to this work with: Harm-Jan Westra, Juha Karjalainen, Daria V. Zhernakova

Affiliation Department of Genetics, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands

Tõnu Esko

Affiliation Institute of Molecular and Cell Biology and Estonian Genome Center, University of Tartu, Tartu, Estonia

Barbara Hrdlickova

Affiliation Department of Genetics, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands

Rodrigo Almeida

Affiliations Department of Genetics, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands, Graduate Program in Health Sciences, University of Brasilia School of Health Sciences, Brasilia, Brazil

Alexandra Zhernakova

Affiliation Department of Genetics, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands

Eva Reinmaa

Affiliation Institute of Molecular and Cell Biology and Estonian Genome Center, University of Tartu, Tartu, Estonia

Urmo Võsa

Affiliation Institute of Molecular and Cell Biology and Estonian Genome Center, University of Tartu, Tartu, Estonia

Marten H. Hofker

Affiliation Molecular Genetics Section, Department of Pathology and Medical Biology, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands

Rudolf S. N. Fehrmann

Affiliation Department of Genetics, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands

Jingyuan Fu

Affiliation Department of Genetics, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands

Sebo Withoff

Affiliation Department of Genetics, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands

Andres Metspalu

Affiliation Institute of Molecular and Cell Biology and Estonian Genome Center, University of Tartu, Tartu, Estonia

Lude Franke

These authors were joint senior authors on this work.

Affiliation Department of Genetics, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands

Cisca Wijmenga

c.wijmenga@umcg.nl

These authors were joint senior authors on this work.

Affiliation Department of Genetics, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: VK LF CW. Performed the experiments: VK H-JW JK DVZ TE BH RA AZ ER UV JF. Analyzed the data: VK H-JW JK DVZ. Contributed reagents/materials/analysis tools: MHH RSNF AM CW. Wrote the paper: VK H-JW SW LF CW.