About the Authors

Ben E. Housden

Current address: Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Affiliation Department of Physiology, Development, and Neuroscience, University of Cambridge, Cambridge, United Kingdom

Audrey Q. Fu

Current address: Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America

Affiliations Department of Physiology, Development, and Neuroscience, University of Cambridge, Cambridge, United Kingdom, Cambridge Systems Biology Centre, University of Cambridge, Cambridge, United Kingdom

Alena Krejci

Current address: University of South Bohemia, Faculty of Science, Ceske Budejovice, Czech Republic

Current address: Biology Centre, Czech Academy of Sciences, Ceske Budejovice, Czech Republic

Affiliation Department of Physiology, Development, and Neuroscience, University of Cambridge, Cambridge, United Kingdom

Fred Bernard

Current address: Institut Pasteur, Department of Developmental Biology, CNRS URA 2578, Paris, France

Affiliation Department of Physiology, Development, and Neuroscience, University of Cambridge, Cambridge, United Kingdom

Bettina Fischer

Affiliations Cambridge Systems Biology Centre, University of Cambridge, Cambridge, United Kingdom, Department of Genetics, University of Cambridge, Cambridge, United Kingdom

Simon Tavaré

Affiliation Department of Applied Mathematics and Theoretical Physics, University of Cambridge, Cambridge, United Kingdom

Steven Russell

Affiliations Cambridge Systems Biology Centre, University of Cambridge, Cambridge, United Kingdom, Department of Genetics, University of Cambridge, Cambridge, United Kingdom

Sarah J. Bray

sjb32@cam.ac.uk

Affiliation Department of Physiology, Development, and Neuroscience, University of Cambridge, Cambridge, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BEH AK FB BF SR SJB. Performed the experiments: BEH AK BF. Analyzed the data: BEH AQF FB BF ST SR SJB. Contributed reagents/materials/analysis tools: AQF FB BF ST SR. Wrote the paper: BEH AQF AK FB SR SJB.