About the Authors

Geeta N. Eick

Affiliations Institute of Ecology and Evolution, University of Oregon, Eugene, Oregon, United States of America, Howard Hughes Medical Institute, Eugene, Oregon, United States of America

Jennifer K. Colucci

Affiliation Biochemistry Department, Emory University School of Medicine, Atlanta, Georgia, United States of America

Michael J. Harms

Affiliation Institute of Ecology and Evolution, University of Oregon, Eugene, Oregon, United States of America

Eric A. Ortlund

Affiliation Biochemistry Department, Emory University School of Medicine, Atlanta, Georgia, United States of America

Joseph W. Thornton

joet1@uchicago.edu

Affiliations Institute of Ecology and Evolution, University of Oregon, Eugene, Oregon, United States of America, Howard Hughes Medical Institute, Eugene, Oregon, United States of America, Department of Human Genetics and Department of Ecology and Evolution, University of Chicago, Chicago, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JWT GNE JKC MJH EAO. Performed the experiments: GNE JKC MJH. Analyzed the data: GNE JKC MJH JWT EAO. Wrote the paper: JWT GNE MJH JKC EAO.