About the Authors

Rebecca E. McIntyre, Martin Del Castillo Velasco-Herrera, Louise van der Weyden, David J. Adams
Experimental Cancer Genetics, Wellcome Trust Sanger Institute, Hinxton, United Kingdom

Pavithra Lakshminarasimhan Chavali, Fanni Gergely
Cancer Research UK Cambridge Research Institute, Li Ka Shing Centre and Department of Oncology, University of Cambridge, Cambridge, United Kingdom

Ozama Ismail, Damian M. Carragher, Sanger Mouse Genetics Project, Ramiro Ramirez-Solis, Jeanne Estabel, Jacqueline K. White
Mouse Genetics Project, Wellcome Trust Sanger Institute, Hinxton, United Kingdom

Gabriela Sanchez-Andrade, Darren W. Logan
Genetics of Instinctive Behaviour, Wellcome Trust Sanger Institute, Hinxton, United Kingdom

Josep V. Forment, Stephen P. Jackson
The Gurdon Institute and Department of Biochemistry, University of Cambridge, Cambridge, United Kingdom

Beiyuan Fu, Fengtang Yang
Molecular Cytogenetics, Wellcome Trust Sanger Institute, Hinxton, United Kingdom

Andrew Edwards
Wellcome Trust Center for Human Genetics, Oxford, United Kingdom

Ferdia A. Gallagher
Department of Radiology, Addenbrooke's Hospital, University of Cambridge, Cambridge, United Kingdom

Mark J. Arends
Department of Pathology, Addenbrooke's Hospital, University of Cambridge, Cambridge, United Kingdom

Stephen H. Tsang, Vinit B. Mahajan
Department of Ophthalmology and Visual Sciences, University of Iowa, Iowa City, Iowa, United States of America

Stephen H. Tsang
Bernard and Shirlee Brown Glaucoma Laboratory, Department of Ophthalmology, College of Physicians and Surgeons, Columbia University, New York, New York, United States of America

Cheryl L. Scudamore
Department of Pathology and Infectious Diseases, Royal Veterinary College, London, United Kingdom

Corresponding Author

da1@sanger.ac.uk

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DJA JKW RR-S REM GS-A OI DMC AE SPJ. Performed the experiments: REM OI DMC GS-A MDCV-H AE VBM PLC JVF BF FY JE LvdW JKW MJA. Analyzed the data: REM OI GS-A MDCV-H AE FAG DWL MJA SHT SMGP CLS FG. Wrote the paper: REM FG SPJ DJA.