About the Authors

Carsten Merkwirth

Address Current address: The Salk Institute for Biological Studies, La Jolla, California, United States of America

Affiliations Institute for Genetics, University of Cologne, Cologne, Germany, Cologne Excellence Cluster on Cellular Stress Responses in Aging-Associated Diseases (CECAD), University of Cologne, Cologne, Germany, Center for Molecular Medicine (CMMC), University of Cologne, Cologne, Germany

Paola Martinelli

Contributed equally to this work with: Paola Martinelli, Anne Korwitz

Affiliation Institute for Zoology, University of Cologne, Cologne, Germany

Anne Korwitz

Contributed equally to this work with: Paola Martinelli, Anne Korwitz

Affiliations Institute for Genetics, University of Cologne, Cologne, Germany, Cologne Excellence Cluster on Cellular Stress Responses in Aging-Associated Diseases (CECAD), University of Cologne, Cologne, Germany, Center for Molecular Medicine (CMMC), University of Cologne, Cologne, Germany

Michela Morbin

Affiliation Neuropathology and Neurology 5, IRCCS Foundation, Neurological Institute Carlo Besta, Milano, Italy

Hella S. Brönneke

Affiliation Cologne Excellence Cluster on Cellular Stress Responses in Aging-Associated Diseases (CECAD), University of Cologne, Cologne, Germany

Sabine D. Jordan

Affiliation Institute for Genetics, University of Cologne, Cologne, Germany

Elena I. Rugarli

Affiliations Cologne Excellence Cluster on Cellular Stress Responses in Aging-Associated Diseases (CECAD), University of Cologne, Cologne, Germany, Center for Molecular Medicine (CMMC), University of Cologne, Cologne, Germany, Institute for Zoology, University of Cologne, Cologne, Germany

Thomas Langer

Thomas.Langer@uni-koeln.de

Affiliations Institute for Genetics, University of Cologne, Cologne, Germany, Cologne Excellence Cluster on Cellular Stress Responses in Aging-Associated Diseases (CECAD), University of Cologne, Cologne, Germany, Center for Molecular Medicine (CMMC), University of Cologne, Cologne, Germany, Max Planck Institute for Biology of Aging, Cologne, Germany

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CM PM AK HSB EIR TL. Performed the experiments: CM PM AK MM HSB SDJ. Analyzed the data: CM PM AK MM HSB SDJ. Wrote the paper: CM EIR TL.