About the Authors

Yoko Nakano

Affiliations Department of Anatomy and Cell Biology, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America, Inflammation Program, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Israt Jahan

Affiliation Department of Biology, College of Liberal Arts and Sciences, University of Iowa, Iowa City, Iowa, United States of America

Gregory Bonde

Affiliation Department of Anatomy and Cell Biology, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Xingshen Sun

Affiliation Department of Anatomy and Cell Biology, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Michael S. Hildebrand

Affiliation Department of Otolaryngology–Head and Neck Surgery, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America

John F. Engelhardt

Affiliation Department of Anatomy and Cell Biology, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Richard J. H. Smith

Affiliation Department of Otolaryngology–Head and Neck Surgery, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Robert A. Cornell

Affiliation Department of Anatomy and Cell Biology, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Bernd Fritzsch

Affiliation Department of Biology, College of Liberal Arts and Sciences, University of Iowa, Iowa City, Iowa, United States of America

Botond Bánfi

botond-banfi@uiowa.edu

Affiliations Department of Anatomy and Cell Biology, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America, Inflammation Program, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America, Department of Otolaryngology–Head and Neck Surgery, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America, Department of Internal Medicine, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YN RAC BF BB. Performed the experiments: YN IJ GB XS MSH. Analyzed the data: YN IJ GB MSH BF JFE RJHS RAC BB. Wrote the paper: YN BB.